Reportedly, in preparation for the role, Ifans did not wash himself or brush his teeth.
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In 2016, Ifans played Fool alongside Glenda Jackson in Deborah Warner's production of King Lear, at The Old Vic.
He returned to The Old Vic to play Ebenezer Scrooge in Matthew Warchus' production of A Christmas Carol (adapted by Jack Thorne) in 2017 and in 2018 returned to the National Theatre to play King Berenger in Patrick Marber's new adaptation of Eugene Ionesco's Exit the King.
Ifans revealed in March 2009 that he was to appear in Harry Potter and the Deathly Hallows – Part 1 (2010).
He played Xenophillius Lovegood, editor of the wizarding magazine The Quibbler and father of the eccentric Luna Lovegood.
Other film roles include: Eyeball Paul in Kevin & Perry Go Large (2000), Nigel in The Replacements (2000), Iki in The 51st State (2001), William Dobbin in Vanity Fair (2004), and Vladis Grutas in Hannibal Rising (2007).